Protologue Description: Jaltomata cajacayensis S. Leiva & Mione, sp. nov. TYPE: Peru. Ancash: Bolognesi, highway from Pativilca to Recuay and Huaraz, km 90, town of Cajacay, 2540 m, open sun, roadside with Agave andina and Carica candicans, 18 Jan. 1988, T. Mione, S. Leiva G. & L. Yacher. 624 (holotype, NY; isotype CONN). Figure 1.

Planta fruticosa; axes juvenes, folia faciesque abaxialis calycis obtecta pilis indivisis apice glandiferentibus; folia ovata vel deltata, ad 9 cm longitudine, 10 cm latitudine, petiolo ad 3.2 longitudine; inflorescentia floribus 14 ut maximum; calyx 13 mm diametro maturitate fructus; corolla brevissime, tuberosa, limbo reflexo 10.3-19.2 mm diametro, olivacea; filamenta villosa secus proximales 4/5 partes longitudinis; bacca aurantiaca, ad 9 mm diametro.

Shrub to 1.1 m high. Young axes, peduncles, pedicels, and abaxial faces of calyx villous with erect, gland-tipped finger hairs to 3 mm long. Woody stems glabrous, terete, to at least 1.5 cm diam. Leaves alternate, often geminate, the blade ovate or deltoid, to 9 X 10 cm, pubescence of gland-tipped finger hairs on both faces, the margin repand, sinuate or dentate with up to 5 pairs of braod, rounded teeth; petiole to 3.2 cm long. Inflorescence axillary or less commonly arising from a stem dichotomy, umbellate, 6- to 14-flowered. Peduncle to 15 mm long, pedicel to 20 mm long. Calyx green at anthesis, 3-4 mm from pedicel to tip of lobe, ciliate with both branchlet hairs to 0.25 mm long and gland-tipped finger hairs to 1 mm long; adaxially pubescent with finger hairs to 0.4 mm long; calyx at fruit maturity ( from Leiva 1739) stellate and 13 mm diam. Corolla short-tubular with a reflexed limb, dull orange green, the tube (obscured by pressing) 1.5-2 X 6.5-8 mm, within the tube two dark green maculae flanking the radial vein extending to the tip of each corolla lobe, narrow purple ring surrounding mouth of tube, the limb 10.3-19.2 mm diam., the 5 lobes alternating with lobules, adaxially glabrous within tube and the limb pilosulose, the hairs 0.21-0.5 mm long, abaxially with finger hairs, some gland tipped, 00.15-1.2 mm long, ciliate with finger hairs to 0.6 mm long. Stalked multicellular glands 75-90 µm long, abundant on both faces of calyx and abaxial corolla, uncommon on adaxial corolla. Stamens 7 mm long; filaments villous on proximal 4/5 of the length, the finger hairs purple, to 1.5 mm long, decreasing in length distally; anthers cream-colored, 1-2.1 mm long. Style 3.5-10 mm long; stigma capitate, dark green and exserted to 5.5 mm beyond anthers on living plants, 0.5-0.9 mm wide, the papillae 45 µm long. Nectar disk broad, girdling base of the ovary (Fig. 1H). Gynoecium glabrous. Mature berries orange (not seen, from informant at type locality), nearly spherical, 5-9 mm diam. Seeds (leiva 1739) numerous, subtriangular to reniform, alveolate, 1.4-1.7 X 1.1-1.3 mm, 0.5-0.55 mm thick.

Paratypes. PERU. Ancash: Bolognesi, Cajacay, 2600 m, 18 Nov. 1995, Leiva 1739 (AAU, CCVS, CORD, F, HAO, HUT, MO, NY, USM), 18 Jan. 1998, Leiva et al. 2132 (F, HAO).




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